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North America yielded its first pterosaur to the prolific palaeontologist O. With an estimated wingspan of 6 m, Pteranodon was huge compared to most earlier discoveries.

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While these discoveries and others were being made, varied opinions on the nature and life style of pterosaurs were appearing, and they were variously depicted as swimming creatures, as bats, marsupials, or as kin of birds.

By the early s, it was generally agreed that pterosaurs were bat-like flying reptiles and, inHarry Seeley — published Dragons of the Air, the first book devoted to pterosaurs. South American Cretaceous pterosaurs have proved to be among the most important in the world, but not until was the first pterosaur from the now famous Santana Formation of Brazil discovered.

Since then a significant number of new kinds from around the world around 70 genera are presently recognized have revealed previously unimagined morphologies and maximum sizes. UntilPteranodon sternbergi Figure 1 Life restoration of the Late Jurassic pterodactyloid, Pterodactylusfrom the German Solnhofen Limestone in a quadrupedal stance.

Note the presence of body hair and the soft tissue head crest. Reproduced with permission from Dino Frey. Geological Society Special Publication London: The Geological Society of London.

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Related pterosaurs of similar or larger size were discovered in the s in Spain and eastern Europe. The Pterosaur Skeleton The pterosaur skeleton was highly modified for flight, and the most obvious features are the huge size of the skull compared with the body and the extreme elongation of one of the fingers. Like birds, most pterosaurs had hollow bones with foramina small openingsindicating that they contained air sacs connected to the lungs.

Pterosaur bones were sup- ported internally by struts, and the bone walls them- selves, usually no thicker than 2 mm, are composed of multiple overlapping layers and thus combine lightness with strength. Pterosaur skull morphology is varied, although the majority had long, slim, shallow jaws and all had large orbits eye-sockets.

In basal pterosaurs, the external nostril was separate from an opening in front of the orbit called the antorbital fenestra. In pterodactyloids, these two openings merged into a single one called the nasoantorbital fenestra.

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Ptero- saur teeth were extremely variable. Widely spaced pointed teeth, were widespread and from ancestors with teeth like these evolved speed ​​dating london clapham with fang-like teeth at the jaw tips and the unique Istiodactylus with its short petal-shaped teeth.

The Late Triassic Eudimorphodon and Austriadactylus possessed multicusped teeth while elongate, slender teeth numbering in the hundreds evolved in the ctenochas- matoids.

Toothlessness evolved several times. Some pterosaurs skulls sport bony crests at the jaw tips, along the midline or at the back of the skull. Figure 2 Wing skeleton of an ornithocheiroid pterodactyloid.

Unlike birds and bats, the main wing spar in ptero- saurs was formed by a hypertrophied digit Figure 2. However, a rod-shaped bone projecting from the pterosaur wrist, called the pteroid bone, has at times been argued to represent the first hand digit. This is a minority view today but, if it is correct, then pterosaurs have five hand digits and the wing finger is the fifth.

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